ATP hydrolysis is required for relocating cohesin from sites occupied by its Scc2/4 loading complex

Bin Hu, Takehiko Itoh, Ajay Mishra, Yuki Katoh, Kok-Lung Chan, William Upcher, Camilla Godlee, Maurici B. Roig, Katsuhiko Shirahige, Kim Nasmyth

Research output: Contribution to journalArticlepeer-review

134 Citations (Scopus)

Abstract

The Cohesin complex that holds sister chromatins together until anaphase is comprised of three core subunits: Smc1 and Smc3, two long-rod-shaped proteins with an ABC-like ATPase head (nucleotide-binding domain [NBD]) and a dimerization domain linked by a 50 nm long intramolecular antiparallel coiled-coil, and Scc1, an α-kleisin subunit interconnecting the NBD domains of Smc1 and Smc3. Cohesin's stable association with chromosomes is thought to involve entrapment of chromatin fibers by its tripartite Smc1-Smc3-Scc1 ring via a poorly understood mechanism dependent on a separate Scc2/4 loading complex. A key issue concerns where entrapment initially takes place: at sites where cohesin is found stably associated or at distinct "loading" sites from which it translocates.
Original languageEnglish
Pages (from-to)12-24
Number of pages12
JournalCurrent Biology
Volume21
Issue number1
DOIs
Publication statusPublished - 11 Jan 2011

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